I really like the anatomy. The improvement is very well visible. The head is amazing, I don't find anything wrong with it. The chest is a good level of fur and fluff. Although the body could be a LITTLE bit shorter. Other then that it's well done. The back legs could do a little work, but I bet that could be fixed on your own time. No rush. As well as the front legs. The tail is not to fluffy it's just perfect. You did a wonderful job. Your future work will look wonder full in time. Keep working hard!
For starters, the atonomy isn't as built as a living wolf's. If your aiming for a cartooned point of view, or realistic, I suppose you look at real wolves.(Which is a good starter) and study their paws meanwhile. Shrink the head and make the body more plump. You might aswell want to shrink the tail and study a wolf's tail to perfect. Try to study the eye, if your trying cartoony or realism. (Eyes are concealed, not open!) Try to sketch more of each body part on a different page then once you feel your more open to parts of a wolf. Nice job anyways!